Adder's-tongue family
Ophioglossaceae

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Adder's-tongue family

Ophioglossaceae

ハナヤスリ科

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One of the most primitive lineages among ferns. Leaves have a distinctive structure where a trophophore and a sporophore differentiate from a single common stalk. Includes Ophioglossum, Botrychium, and related genera, with multiple species native to Japan.

Key Characteristics

  • Trophophore and sporophore differentiate from a single leaf (common stalk)
  • Sporangia thick-walled (eusporangiate), lacking an annulus
  • Prothallus subterranean with mycorrhizal association
  • Some species have extremely high chromosome numbers (among the highest known)

Phylogenetic Position

Vascular plants > Polypodiophyta > Ophioglossopsida > Ophioglossales > Ophioglossaceae

Divergence Era

Devonian to Carboniferous (approx. 360 million years ago onward)

Representative Genera

ハナヤスリ属(Ophioglossum)ナツノハナワラビ属(Botrychium)

Related Evolution Events

  • Retention of eusporangiate sporangia (a primitive characteristic)
  • Extreme polyploidization (chromosome numbers reaching 2n=1440 in some species)
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Plants in Adder's-tongue family on this site

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📚くらべてわかるシダ(山と渓谷社)

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