Buckthorn family
Rhamnaceae

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Buckthorn family

Rhamnaceae

クロウメモドキ科

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A family of about 52 genera and 900 species including Rhamnus, Berchemia, Ziziphus, and Hovenia. Widely distributed from the tropics to temperate regions worldwide. About 10 genera are native to Japan, where the sweet fleshy peduncle of Hovenia dulcis is edible in autumn.

Key Characteristics

  • Predominantly woody (trees, shrubs, or woody climbers)
  • Flowers small, pentamerous; petals small and concave, enclosing the stamens
  • Many species thorny
  • Fruit types diverse: drupes, samaras, or capsules
  • Leaves alternate, often with 3 prominent main veins

Morphological Traits

A family may include species with different trait values — multiple values indicate the range within the family.

Leaf arrangement

Alternate / Opposite

Leaf type

Simple

Venation

Pinnate / Palmate

Leaf margin

Entire

Growth form

Shrub / Tree / Vine / Herb

Evergreen/Deciduous

Deciduous

Leaf dissection

Undivided

Stipules

Present / Absent

Flower symmetry

Actinomorphic

Petal count

5 petals / Many

Petal fusion

Fused

Ovary position

Superior / Inferior / Half-inferior

Stamen count

3-4

Plant sex

Dioecious / Hermaphrodite / Monoecious

Phylogenetic Position

Angiosperms > Eudicots > Core eudicots > Rosids > Rosales > Rhamnaceae

Divergence Era

Late Cretaceous (approx. 80 million years ago)

Representative Genera

クロウメモドキ属(Rhamnus)ケンポナシ属(Hovenia)クマヤナギ属(Berchemia)ナツメ属(Ziziphus)

Related Evolution Events

  • Widespread distribution from tropics to temperate regions with evolution of diverse fruit types
  • Acquisition of nitrogen fixation through Frankia symbiosis in some lineages
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Plants in Buckthorn family on this site

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